Nikhil Milind

PhD Candidate

Stanford University

About Me

I am a PhD student in Jonathan Pritchard's lab at Stanford University. I am broadly interested in statistical and population genetics.

I have recently been thinking about the relationship between gene dosage and complex traits. We introduced the concept of the gene dosage response curve, which is a conceptual tool to think about these relationships using population-level or experimental data.

I was previously a Master's student with Emma Davenport, where I worked on using functional genomics to better understand sepsis. As an undergraduate student, I worked with David Aylor on mouse quantitative genetics and Gregory Carter on statistical genetics approaches in Alzheimer's disease cohorts.

Education

M.Phil. in Biological Sciences

University of Cambridge | Wellcome Sanger Institute (2022)

B.S. in Genetics and Computer Science

North Carolina State University (2021)

Recent Publications

Year Title
2025 Causal modelling of gene effects from regulators to programs to traits
2025 Allele frequencies at recessive disease genes are mainly determined by pleiotropic effects in heterozygotes
2025 Specificity, length and luck drive gene rankings in association studies
2024 Buffering and non-monotonic behavior of gene dosage response curves for human complex traits
2024 eQTLs identify regulatory networks and drivers of variation in the individual response to sepsis

Recent Posts

Year Title
2025 Numerical gradients through numerical integrals